Water in the liquid phase, the biogenic elements, and energy are the fundamental requirements for habitability. But are there additional environmental constraints? We consider this in the next section.
Grand prismatic spring in yellowstone national park.
Life in extreme conditions
At a chemical level, life consists of many types of molecules that interact with one another to carry out the processes of life. In addition to water, elemental raw materials, and energy, life also needs an environment in which those complicated molecules are stable (don’t break down before they can do their jobs) and their interactions are possible. Your own biochemistry works properly only within a very narrow range of about 10 °C in body temperature and two-tenths of a unit in blood pH (pH is a numerical measure of acidity, or the amount of free hydrogen ions). Beyond those limits, you are in serious danger.
Life overall must also have limits to the conditions in which it can properly work but, as we will see, they are much broader than human limits. The resources that fuel life are distributed across a very wide range of conditions. For example, there is abundant chemical energy to be had in hot springs that are essentially boiling acid (see [link] ). This provides ample incentive for evolution to fill as much of that range with life as is biochemically possible. An organism (usually a microbe) that tolerates or even thrives under conditions that most of the life around us would consider hostile, such as very high or low temperature or acidity, is known as an extremophile (where the suffix -phile means “lover of”). Let’s have a look at some of the conditions that can challenge life and the organisms that have managed to carve out a niche at the far reaches of possibility.
Both high and low temperatures can cause a problem for life. As a large organism, you are able to maintain an almost constant body temperature whether it is colder or warmer in the environment around you. But this is not possible at the tiny size of microorganisms; whatever the temperature in the outside world is also the temperature of the microbe, and its biochemistry must be able to function at that temperature. High temperatures are the enemy of complexity—increasing thermal energy tends to break apart big molecules into smaller and smaller bits, and life needs to stabilize the molecules with stronger bonds and special proteins. But this approach has its limits.